I can’t quote from the article, but on page 2 it says: “nicotine exposure could have a subtle beneficial effect on the host by limiting the growth of certain respiratory tract and eneric pathogens, as the enter the body trough the oral and nasal passages”
The pathogens are a variety of bacteria and the fungi c. albicans and crypt. neoformans.
And this is the nicotine only. I will look more into it, but I’m a bit fascinated about killing the fungi in my lungs so I don’t get so sick when the weather get more moist.
Note: tobacco mostly contains some molds and fungi too.
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